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To determine the subcellular localization and expression pattern of VPS-18, we analyzed a VPS-18::GFP fusion protein under the control of the vps-18 promoter (Pvps-18vps-18::gfp). VPS-18::GFP was ubiquitously expressed in early embryo and seemed cytoplasmic ( Figure 4 A, a and b). In early larvae, VPS-18::GFP was mainly seen in hypodermal cells, seam cells and some body wall muscle cells ( Figure 4 A, c and d; data not shown). In adult hermaphrodite, VPS-18::GFP was detected in coelomocytes ( Figure 4 A, e and f), the C. elegans scavenger cells functioning similarly to mammalian macrophages for endocytosing fluid from pseudocoelom ( Fares and Greenwald, 2001a ). VPS-18::GFP signal was also present in gonadal sheath cells ( Figure 4 A, g-j). In addition, VPS-18::GFP was expressed in intestine and spermatheca (data not shown).